:root {
    --forest-900: #0B2A20;
    --forest-800: #123024;
    --forest-700: #1B4332;
    --forest-600: #2D6A4F;
    --moss-400: #6E9884;
    --moss-200: #CBDED2;
    --mist: #F4F8F5;
    --paper: #FFFFFF;
    --ink: #16241C;
    --ink-muted: #57685D;
    --border: #DEE8E1;

    --font-display: "Fraunces", Georgia, serif;
    --font-body: "Inter", "Segoe UI", sans-serif;
    --font-mono: "IBM Plex Mono", ui-monospace, monospace;
}

* {
    box-sizing: border-box;
}

html {
    scroll-behavior: smooth;
}

body {
    margin: 0;
    background: var(--paper);
    color: var(--ink);
    font-family: var(--font-body);
    line-height: 1.5;
}

h1, h2, h3 {
    font-family: var(--font-display);
    color: var(--forest-800);
    font-weight: 600;
    margin: 0;
}

p {
    margin: 0;
}

a {
    color: inherit;
}

.container {
    width: 100%;
    max-width: 1180px;
    margin: auto;
    padding: 0 24px;
}

.eyebrow {
    display: inline-block;
    font-family: var(--font-mono);
    font-size: 0.78rem;
    letter-spacing: 0.14em;
    text-transform: uppercase;
    color: var(--forest-600);
    margin-bottom: 14px;
}

/* =====================
   NAVBAR
===================== */

.navbar-custom {
    position: sticky;
    top: 0;
    z-index: 1000;
    background: rgba(255, 255, 255, .92);
    backdrop-filter: blur(10px);
    border-bottom: 1px solid var(--border);
}

.navbar-content {
    height: 76px;
    display: flex;
    align-items: center;
    justify-content: space-between;
}

.navbar-brand {
    display: flex;
    align-items: center;
    gap: 10px;
    text-decoration: none;
    color: var(--forest-800);
    font-family: var(--font-display);
    font-weight: 600;
    font-size: 1.2rem;
}

.navbar-logo {
    width: 32px;
    height: 32px;
    object-fit: contain;
}

.navbar-links {
    display: flex;
    gap: 30px;
}

.navbar-links a {
    text-decoration: none;
    color: var(--ink-muted);
    font-weight: 500;
    font-size: 0.95rem;
    transition: .2s;
}

.navbar-links a:hover {
    color: var(--forest-700);
}

.nav-cta {
    background: var(--forest-700);
    color: white;
    text-decoration: none;
    padding: 11px 22px;
    border-radius: 999px;
    font-weight: 600;
    font-size: 0.92rem;
    transition: .2s;
}

.nav-cta:hover {
    background: var(--forest-800);
}

/* =====================
   HERO
===================== */

.hero {
    position: relative;
    padding: 76px 0 0;
    overflow: hidden;
}

.hero-texture {
    position: absolute;
    inset: 0;
    background-image: url("img/topo-pattern.svg");
    background-repeat: repeat;
    background-size: 480px;
    pointer-events: none;
}

.hero-content {
    position: relative;
    display: flex;
    align-items: center;
    justify-content: space-between;
    gap: 60px;
    padding-bottom: 70px;
}

.hero-text {
    flex: 1.1;
}

.hero-eyebrow {
    display: inline-block;
    font-family: var(--font-mono);
    font-size: 0.8rem;
    letter-spacing: 0.1em;
    text-transform: uppercase;
    color: var(--forest-600);
    background: var(--mist);
    border: 1px solid var(--border);
    padding: 8px 16px;
    border-radius: 999px;
    margin-bottom: 26px;
}

.hero h1 {
    font-size: 3.6rem;
    line-height: 1.08;
    font-weight: 600;
    max-width: 620px;
    color: var(--forest-800);
    margin-bottom: 22px;
}

.hero p {
    max-width: 520px;
    font-size: 1.12rem;
    line-height: 1.75;
    color: var(--ink-muted);
}

.hero-actions {
    display: flex;
    gap: 16px;
    margin-top: 34px;
    flex-wrap: wrap;
}

.btn-primary-custom {
    background: var(--forest-700);
    color: white;
    text-decoration: none;
    padding: 15px 26px;
    border-radius: 10px;
    font-weight: 600;
    border: 1px solid var(--forest-700);
    transition: .25s;
    display: inline-block;
    cursor: pointer;
    font-size: 0.98rem;
    font-family: var(--font-body);
}

.btn-primary-custom:hover {
    background: var(--forest-800);
    border-color: var(--forest-800);
}

.btn-outline-custom {
    border: 1px solid var(--border);
    color: var(--forest-800);
    text-decoration: none;
    padding: 15px 26px;
    border-radius: 10px;
    font-weight: 600;
    transition: .2s;
}

.btn-outline-custom:hover {
    border-color: var(--forest-600);
    background: var(--mist);
}

.hero-visual {
    flex: 0.9;
    display: flex;
    justify-content: center;
}

.hero-visual-frame {
    position: relative;
    width: 100%;
    max-width: 380px;
    aspect-ratio: 1 / 1;
    background: var(--mist);
    border: 1px solid var(--border);
    border-radius: 28px;
    display: flex;
    align-items: center;
    justify-content: center;
}

.hero-frog-mark {
    width: 62%;
    opacity: 0.9;
}

.field-card {
    position: absolute;
    background: white;
    border-radius: 16px;
    padding: 14px 18px;
    box-shadow: 0 14px 30px rgba(11, 42, 32, .12);
    font-size: 0.85rem;
    display: flex;
    flex-direction: column;
    gap: 2px;
}

.field-card strong {
    color: var(--forest-800);
    font-size: 0.92rem;
}

.field-card span {
    color: var(--ink-muted);
}

.field-card small {
    color: var(--moss-400);
    font-family: var(--font-mono);
    font-size: 0.72rem;
}

.field-card--status {
    top: 18px;
    left: -18px;
    flex-direction: row;
    align-items: center;
    gap: 10px;
    min-width: 190px;
}

.pulse-dot {
    width: 10px;
    height: 10px;
    border-radius: 50%;
    background: var(--forest-600);
    box-shadow: 0 0 0 4px rgba(45, 106, 79, .18);
    flex-shrink: 0;
}

.field-card--coords {
    bottom: 22px;
    right: -22px;
    min-width: 170px;
}

.stats-row {
    position: relative;
    display: grid;
    grid-template-columns: repeat(4, 1fr);
    border-top: 1px solid var(--border);
    padding: 34px 24px;
}

.stat {
    display: flex;
    flex-direction: column;
    gap: 4px;
    text-align: center;
    border-left: 1px solid var(--border);
    padding: 0 12px;
}

.stat:first-child {
    border-left: none;
}

.stat-number {
    font-family: var(--font-display);
    font-size: 2.4rem;
    font-weight: 600;
    color: var(--forest-700);
}

.stat-number--sm {
    font-size: 2rem;
}

.stat-label {
    font-size: 0.85rem;
    color: var(--ink-muted);
}

/* =====================
   SECTION HEADERS
===================== */

.section-header {
    max-width: 640px;
    margin-bottom: 56px;
}

.section-header--split {
    display: flex;
    justify-content: space-between;
    align-items: flex-end;
    max-width: none;
}

.section-header h2 {
    font-size: 2.3rem;
    line-height: 1.2;
    margin-bottom: 14px;
}

.section-header p {
    color: var(--ink-muted);
    line-height: 1.75;
}

/* =====================
   ABOUT
===================== */

.about {
    padding: 100px 0;
    background: var(--mist);
}

.about-grid {
    display: grid;
    grid-template-columns: 1.5fr 1fr;
    gap: 70px;
}

.about-intro h2 {
    font-size: 2.3rem;
    margin-bottom: 22px;
    line-height: 1.25;
}

.about-intro p {
    color: var(--ink-muted);
    line-height: 1.8;
    margin-bottom: 18px;
}

.pillars {
    list-style: none;
    margin: 0 0 40px;
    padding: 0;
    border-top: 1px solid var(--border);
}

.pillars li {
    display: flex;
    align-items: baseline;
    justify-content: space-between;
    padding: 18px 0;
    border-bottom: 1px solid var(--border);
}

.pillars em {
    font-family: var(--font-display);
    font-style: italic;
    font-size: 1.5rem;
    color: var(--forest-800);
}

.pillars span {
    color: var(--ink-muted);
    font-size: 0.9rem;
}

.founding-note {
    display: flex;
    flex-direction: column;
    gap: 4px;
    background: var(--paper);
    border: 1px solid var(--border);
    border-radius: 16px;
    padding: 22px 24px;
}

/* =====================
   AREAS
===================== */

.areas {
    padding: 100px 0;
}

.areas-grid {
    display: grid;
    grid-template-columns: repeat(4, 1fr);
    gap: 22px;
}

.area-card {
    background: var(--paper);
    border: 1px solid var(--border);
    border-radius: 18px;
    padding: 28px 24px;
    transition: .25s;
}

.area-card:hover {
    transform: translateY(-4px);
    border-color: var(--moss-400);
}

.area-icon {
    width: 46px;
    height: 46px;
    display: flex;
    align-items: center;
    justify-content: center;
    background: var(--mist);
    border: 1px solid var(--border);
    color: var(--forest-700);
    border-radius: 12px;
    font-size: 1.2rem;
    margin-bottom: 18px;
}

.area-card h3 {
    font-size: 1.05rem;
    margin-bottom: 10px;
}

.area-card p {
    color: var(--ink-muted);
    line-height: 1.65;
    font-size: 0.92rem;
}

/* =====================
   TAGS
===================== */

.tag {
    display: inline-block;
    font-family: var(--font-mono);
    font-size: 0.72rem;
    letter-spacing: 0.06em;
    text-transform: uppercase;
    padding: 5px 12px;
    border-radius: 999px;
}

.tag--active {
    background: var(--forest-700);
    color: white;
}

.tag--progress {
    background: var(--moss-200);
    color: var(--forest-800);
}

.tag--planning {
    background: var(--mist);
    color: var(--ink-muted);
    border: 1px solid var(--border);
}

.tag--danger {
    background: var(--forest-800);
    color: white;
}

.tag--warning {
    background: transparent;
    color: var(--forest-700);
    border: 1px solid var(--forest-700);
}

.tag--outline {
    background: transparent;
    color: var(--forest-700);
    border: 1px solid var(--border);
}

/* =====================
   PROJECTS
===================== */

.projects {
    padding: 100px 0;
    background: var(--mist);
}

.projects-grid {
    display: grid;
    grid-template-columns: repeat(2, 1fr);
    gap: 22px;
}

.project-card {
    background: var(--paper);
    border: 1px solid var(--border);
    border-radius: 18px;
    padding: 30px;
}

.project-meta {
    display: flex;
    align-items: center;
    justify-content: space-between;
    margin-bottom: 18px;
}

.project-period {
    font-family: var(--font-mono);
    font-size: 0.8rem;
    color: var(--ink-muted);
}

.project-card h3 {
    font-size: 1.25rem;
    margin-bottom: 6px;
}

.project-ecosystem {
    display: block;
    color: var(--moss-400);
    font-size: 0.85rem;
    font-weight: 600;
    margin-bottom: 14px;
}

.project-card p {
    color: var(--ink-muted);
    line-height: 1.7;
    margin-bottom: 18px;
}

.project-location {
    font-size: 0.85rem;
    color: var(--ink-muted);
}

/* =====================
   SPECIES
===================== */

.species {
    padding: 100px 0;
}

.species-grid {
    display: grid;
    grid-template-columns: repeat(2, 1fr);
    gap: 22px;
    margin-bottom: 36px;
}

.species-card {
    border: 1px solid var(--border);
    border-radius: 18px;
    padding: 30px;
    background: var(--paper);
}

.species-card-head {
    margin-bottom: 16px;
}

.species-card h3 {
    font-size: 1.4rem;
    margin-bottom: 4px;
}

.scientific-name {
    font-style: italic;
    font-family: var(--font-display);
    color: var(--forest-600);
    margin-bottom: 16px;
}

.species-card > p:not(.scientific-name) {
    color: var(--ink-muted);
    line-height: 1.75;
    margin-bottom: 22px;
}

.species-habitat {
    display: flex;
    justify-content: space-between;
    align-items: center;
    border-top: 1px solid var(--border);
    padding-top: 16px;
    font-size: 0.9rem;
}

.species-habitat span {
    color: var(--ink-muted);
}

.species-habitat strong {
    color: var(--forest-800);
}

.link-arrow {
    display: inline-block;
    text-decoration: none;
    color: var(--forest-700);
    font-weight: 600;
    border-bottom: 1px solid var(--forest-700);
    padding-bottom: 2px;
}

/* =====================
   RESEARCH
===================== */

.research {
    padding: 100px 0;
    background: var(--forest-800);
    color: white;
}

.research-grid {
    display: grid;
    grid-template-columns: 0.9fr 1.4fr;
    gap: 70px;
}

.research .eyebrow {
    color: var(--moss-200);
}

.research-intro h2 {
    color: white;
    font-size: 2.1rem;
    margin-bottom: 18px;
    line-height: 1.25;
}

.research-intro p {
    color: rgba(255, 255, 255, .72);
    line-height: 1.8;
    margin-bottom: 26px;
}

.research-links {
    display: flex;
    gap: 26px;
}

.research-links a {
    color: white;
    text-decoration: none;
    font-weight: 600;
    border-bottom: 1px solid rgba(255, 255, 255, .4);
    padding-bottom: 3px;
}

.publication-list {
    list-style: none;
    margin: 0;
    padding: 0;
}

.publication-list li {
    display: flex;
    gap: 26px;
    padding: 28px 0;
    border-bottom: 1px solid rgba(255, 255, 255, .14);
}

.publication-list li:first-child {
    padding-top: 0;
}

.pub-year {
    font-family: var(--font-mono);
    color: var(--moss-200);
    font-size: 0.95rem;
    flex-shrink: 0;
    width: 48px;
}

.pub-type {
    display: block;
    font-family: var(--font-mono);
    font-size: 0.72rem;
    text-transform: uppercase;
    letter-spacing: 0.08em;
    color: var(--moss-200);
    margin-bottom: 8px;
}

.pub-body h3 {
    color: white;
    font-size: 1.1rem;
    line-height: 1.5;
    margin-bottom: 10px;
    font-weight: 500;
}

.pub-authors {
    color: rgba(255, 255, 255, .75);
    font-size: 0.9rem;
    margin-bottom: 4px;
}

.pub-journal {
    color: rgba(255, 255, 255, .5);
    font-size: 0.85rem;
    font-style: italic;
}

/* =====================
   EDUCATION
===================== */

.education {
    padding: 100px 0;
}

.education-grid {
    display: grid;
    grid-template-columns: repeat(3, 1fr);
    gap: 22px;
}

.education-card {
    border: 1px solid var(--border);
    border-radius: 18px;
    padding: 28px;
    transition: .25s;
}

.education-card:hover {
    border-color: var(--moss-400);
    transform: translateY(-4px);
}

.education-meta {
    display: flex;
    align-items: center;
    justify-content: space-between;
    margin-bottom: 20px;
}

.education-category {
    font-family: var(--font-mono);
    font-size: 0.75rem;
    color: var(--ink-muted);
    text-transform: uppercase;
    letter-spacing: 0.06em;
}

.education-card h3 {
    font-size: 1.1rem;
    line-height: 1.5;
    font-weight: 500;
}

/* =====================
   QUOTE
===================== */

.quote-section {
    background: var(--forest-700);
    padding: 90px 0;
}

.quote-section blockquote {
    margin: 0;
    text-align: center;
    color: white;
    font-family: var(--font-display);
    font-style: italic;
    font-weight: 400;
    font-size: 2rem;
    line-height: 1.5;
    max-width: 780px;
    margin: auto;
}

/* =====================
   CONTACT
===================== */

.contact {
    padding: 100px 0;
}

.contact-grid {
    display: grid;
    grid-template-columns: 1fr 1.3fr;
    gap: 60px;
    margin-bottom: 90px;
}

.contact-channels {
    display: flex;
    flex-direction: column;
    gap: 26px;
}

.contact-channel {
    border-top: 1px solid var(--border);
    padding-top: 16px;
    display: flex;
    flex-direction: column;
    gap: 6px;
}

.channel-label {
    font-family: var(--font-mono);
    font-size: 0.78rem;
    text-transform: uppercase;
    letter-spacing: 0.06em;
    color: var(--ink-muted);
}

.contact-channel a {
    text-decoration: none;
    color: var(--forest-800);
    font-weight: 600;
    font-size: 1.05rem;
}

.contact-channel a:hover {
    color: var(--forest-600);
}

.contact-form {
    display: flex;
    flex-direction: column;
    gap: 18px;
    background: var(--mist);
    border: 1px solid var(--border);
    border-radius: 18px;
    padding: 30px;
}

.contact-form label {
    display: flex;
    flex-direction: column;
    gap: 8px;
    font-size: 0.88rem;
    font-weight: 600;
    color: var(--forest-800);
}

.contact-form input,
.contact-form textarea {
    font-family: var(--font-body);
    font-size: 0.95rem;
    padding: 12px 14px;
    border-radius: 10px;
    border: 1px solid var(--border);
    background: white;
    color: var(--ink);
    resize: vertical;
}

.contact-form input:focus,
.contact-form textarea:focus {
    outline: 2px solid var(--forest-600);
    outline-offset: 1px;
}

.contact-form button {
    align-self: flex-start;
    border: none;
}

.social-block {
    border-top: 1px solid var(--border);
    padding-top: 60px;
}

.social-block h3 {
    font-size: 1.6rem;
    margin: 0 0 32px;
}

.social-grid {
    display: grid;
    grid-template-columns: repeat(4, 1fr);
    gap: 18px;
}

.social-card {
    text-decoration: none;
    color: var(--ink);
    border: 1px solid var(--border);
    border-radius: 16px;
    padding: 22px;
    display: flex;
    flex-direction: column;
    gap: 4px;
    transition: .2s;
}

.social-card:hover {
    border-color: var(--forest-600);
    background: var(--mist);
}

.social-card strong {
    color: var(--forest-800);
}

.social-card span {
    font-size: 0.88rem;
    color: var(--ink-muted);
}

.social-card small {
    font-size: 0.78rem;
    color: var(--moss-400);
    margin-top: 6px;
}

/* =====================
   FOOTER
===================== */

.footer-custom {
    background: var(--forest-900);
    color: rgba(255, 255, 255, .7);
    padding: 70px 0 28px;
}

.footer-grid {
    display: grid;
    grid-template-columns: 1.6fr 1fr 1fr 1fr;
    gap: 40px;
    padding-bottom: 50px;
    border-bottom: 1px solid rgba(255, 255, 255, .12);
}

.footer-brand {
    display: flex;
    align-items: center;
    gap: 10px;
    font-family: var(--font-display);
    font-weight: 600;
    color: white;
    font-size: 1.1rem;
    margin-bottom: 16px;
}

.footer-logo {
    width: 26px;
    height: 26px;
}

.footer-brand-block p {
    color: rgba(255, 255, 255, .55);
    line-height: 1.7;
    font-size: 0.92rem;
    margin-bottom: 18px;
    max-width: 320px;
}

.footer-tagline {
    font-family: var(--font-mono);
    font-size: 0.78rem;
    letter-spacing: 0.06em;
    color: var(--moss-200);
}

.footer-col h4 {
    color: white;
    font-family: var(--font-body);
    font-size: 0.85rem;
    text-transform: uppercase;
    letter-spacing: 0.06em;
    margin-bottom: 18px;
    font-weight: 600;
}

.footer-col {
    display: flex;
    flex-direction: column;
    gap: 12px;
}

.footer-col a {
    text-decoration: none;
    color: rgba(255, 255, 255, .6);
    font-size: 0.92rem;
    transition: .2s;
}

.footer-col a:hover {
    color: white;
}

.footer-bottom {
    padding-top: 24px;
    font-size: 0.85rem;
    color: rgba(255, 255, 255, .45);
}

/* =====================
   RESPONSIVE
===================== */

@media (max-width: 992px) {

    .hero-content {
        flex-direction: column;
    }

    .hero-visual {
        width: 100%;
    }

    .about-grid,
    .research-grid,
    .contact-grid {
        grid-template-columns: 1fr;
        gap: 40px;
    }

    .areas-grid {
        grid-template-columns: repeat(2, 1fr);
    }

    .projects-grid,
    .species-grid,
    .education-grid {
        grid-template-columns: repeat(2, 1fr);
    }

    .social-grid {
        grid-template-columns: repeat(2, 1fr);
    }

    .footer-grid {
        grid-template-columns: repeat(2, 1fr);
    }

    .hero h1 {
        font-size: 2.8rem;
    }
}

@media (max-width: 768px) {

    .navbar-links {
        display: none;
    }

    .stats-row {
        grid-template-columns: repeat(2, 1fr);
        row-gap: 24px;
    }

    .stat:nth-child(3) {
        border-left: none;
    }
}

@media (max-width: 640px) {

    .areas-grid,
    .projects-grid,
    .species-grid,
    .education-grid,
    .social-grid {
        grid-template-columns: 1fr;
    }

    .hero h1 {
        font-size: 2.3rem;
    }

    .section-header h2 {
        font-size: 1.8rem;
    }

    .field-card {
        display: none;
    }

    .footer-grid {
        grid-template-columns: 1fr;
    }

    .quote-section blockquote {
        font-size: 1.5rem;
    }
}